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Rice Science ›› 2026, Vol. 33 ›› Issue (1): 113-128.DOI: 10.1016/j.rsci.2025.10.004

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  • 收稿日期:2025-06-27 接受日期:2025-09-01 出版日期:2026-01-28 发布日期:2026-02-03

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. [J]. Rice Science, 2026, 33(1): 113-128.

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链接本文: http://www.ricesci.org/CN/10.1016/j.rsci.2025.10.004

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图/表 7

Table 1. Effects of planting density on yield and yield components of japonica rice.
Year Variety Treatment No. of panicles per hm2 (×104) Total spikelet number
per m2
Seed-setting rate (%) 1000-grain weight
(g)
Yield
(t/hm2)
2023 Ningxiangjing 9 D1 286.11 ± 3.54 d 52 321.5 ± 864.6 c 84.87 ± 0.73 a 24.50 ± 0.20 a 10.73 ± 0.09 c
D2 305.55 ± 2.97 c 55 633.4 ± 355.2 b 84.70 ± 1.01 a 24.44 ± 0.29 a 11.36 ± 0.13 b
D3 322.22 ± 6.00 b 58 193.5 ± 791.1 a 84.11 ± 0.09 a 24.33 ± 0.31 a 11.68 ± 0.12 a
D4 334.44 ± 6.85 a 59 666.2 ± 697.9 a 83.82 ± 0.93 a 24.17 ± 0.15 a 11.91 ± 0.06 a
Mean 312.08 56 453.6 84.37 24.36 11.42
Songxiangjing 1018 D1 281.95 ± 2.60 d 48 225.5 ± 411.3 d 88.39 ± 0.52 a 27.91 ± 0.11 a 11.72 ± 0.04 c
D2 296.83 ± 2.97 c 50 144.7 ± 309.6 c 88.13 ± 0.24 a 27.84 ± 0.08 a 12.20 ± 0.06 b
D3 318.52 ± 3.46 b 52 748.0 ± 238.0 b 87.67 ± 0.28 a 27.76 ± 0.17 a 12.58 ± 0.18 a
D4 332.22 ± 5.66 a 54 099.7 ± 498.9 a 87.36 ± 0.58 a 27.63 ± 0.26 a 12.76 ± 0.09 a
Mean 307.38 51 304.5 87.89 27.79 12.31
2024 Ningxiangjing 9 D1 301.39 ± 2.60 d 52 931.9 ± 442.5 d 84.74 ± 0.35 a 24.32 ± 0.02 a 10.62 ± 0.01 d
D2 317.46 ± 2.97 c 55 003.9 ± 542.1 c 84.61 ± 0.20 a 24.25 ± 0.04 b 11.07 ± 0.02 c
D3 335.19 ± 3.46 b 57 330.3 ± 586.1 b 84.23 ± 0.21 a 24.20 ± 0.03 bc 11.44 ± 0.04 b
D4 355.56 ± 4.16 a 59 580.6 ± 1 251.1 a 83.66 ± 1.19 a 24.15 ± 0.03 c 11.70 ± 0.09 a
Mean 327.40 56 211.7 84.31 24.23 11.21
Songxiangjing 1018 D1 283.33 ± 1.70 d 46 997.6 ± 207.7 d 87.62 ± 0.31 a 27.71 ± 0.02 a 11.23 ± 0.09 d
D2 300.00 ± 1.94 c 48 918.2 ± 226.6 c 87.44 ± 0.20 a 27.67 ± 0.01 ab 11.71 ± 0.06 c
D3 321.30 ± 4.72 b 51 559.4 ± 757.8 b 87.17 ± 0.17 a 27.64 ± 0.02 b 12.21 ± 0.08 b
D4 341.11 ± 6.85 a 54 048.7 ± 1 132.4 a 86.53 ± 0.17 b 27.59 ± 0.03 c 12.62 ± 0.04 a
Mean 311.44 50 381.0 87.19 27.65 11.94
Year (Y) ** NS NS NS **
Variety (V) ** ** ** ** **
Treatment (T) ** ** ** NS **
Y × V ** NS NS NS NS
Y × T NS NS NS NS NS
V × T NS NS NS NS NS
Y × V × T NS NS NS NS NS

Table 1. Effects of planting density on yield and yield components of japonica rice.

Year Variety Treatment No. of panicles per hm2 (×104) Total spikelet number
per m2
Seed-setting rate (%) 1000-grain weight
(g)
Yield
(t/hm2)
2023 Ningxiangjing 9 D1 286.11 ± 3.54 d 52 321.5 ± 864.6 c 84.87 ± 0.73 a 24.50 ± 0.20 a 10.73 ± 0.09 c
D2 305.55 ± 2.97 c 55 633.4 ± 355.2 b 84.70 ± 1.01 a 24.44 ± 0.29 a 11.36 ± 0.13 b
D3 322.22 ± 6.00 b 58 193.5 ± 791.1 a 84.11 ± 0.09 a 24.33 ± 0.31 a 11.68 ± 0.12 a
D4 334.44 ± 6.85 a 59 666.2 ± 697.9 a 83.82 ± 0.93 a 24.17 ± 0.15 a 11.91 ± 0.06 a
Mean 312.08 56 453.6 84.37 24.36 11.42
Songxiangjing 1018 D1 281.95 ± 2.60 d 48 225.5 ± 411.3 d 88.39 ± 0.52 a 27.91 ± 0.11 a 11.72 ± 0.04 c
D2 296.83 ± 2.97 c 50 144.7 ± 309.6 c 88.13 ± 0.24 a 27.84 ± 0.08 a 12.20 ± 0.06 b
D3 318.52 ± 3.46 b 52 748.0 ± 238.0 b 87.67 ± 0.28 a 27.76 ± 0.17 a 12.58 ± 0.18 a
D4 332.22 ± 5.66 a 54 099.7 ± 498.9 a 87.36 ± 0.58 a 27.63 ± 0.26 a 12.76 ± 0.09 a
Mean 307.38 51 304.5 87.89 27.79 12.31
2024 Ningxiangjing 9 D1 301.39 ± 2.60 d 52 931.9 ± 442.5 d 84.74 ± 0.35 a 24.32 ± 0.02 a 10.62 ± 0.01 d
D2 317.46 ± 2.97 c 55 003.9 ± 542.1 c 84.61 ± 0.20 a 24.25 ± 0.04 b 11.07 ± 0.02 c
D3 335.19 ± 3.46 b 57 330.3 ± 586.1 b 84.23 ± 0.21 a 24.20 ± 0.03 bc 11.44 ± 0.04 b
D4 355.56 ± 4.16 a 59 580.6 ± 1 251.1 a 83.66 ± 1.19 a 24.15 ± 0.03 c 11.70 ± 0.09 a
Mean 327.40 56 211.7 84.31 24.23 11.21
Songxiangjing 1018 D1 283.33 ± 1.70 d 46 997.6 ± 207.7 d 87.62 ± 0.31 a 27.71 ± 0.02 a 11.23 ± 0.09 d
D2 300.00 ± 1.94 c 48 918.2 ± 226.6 c 87.44 ± 0.20 a 27.67 ± 0.01 ab 11.71 ± 0.06 c
D3 321.30 ± 4.72 b 51 559.4 ± 757.8 b 87.17 ± 0.17 a 27.64 ± 0.02 b 12.21 ± 0.08 b
D4 341.11 ± 6.85 a 54 048.7 ± 1 132.4 a 86.53 ± 0.17 b 27.59 ± 0.03 c 12.62 ± 0.04 a
Mean 311.44 50 381.0 87.19 27.65 11.94
Year (Y) ** NS NS NS **
Variety (V) ** ** ** ** **
Treatment (T) ** ** ** NS **
Y × V ** NS NS NS NS
Y × T NS NS NS NS NS
V × T NS NS NS NS NS
Y × V × T NS NS NS NS NS
Table 2. Effects of planting density on dry matter accumulation and harvest index of japonica rice.
Year Variety Treatment Dry matter (t/hm2) Harvest index
(%)
Jointing stage Heading stage Maturity stage
2023 Ningxiangjing 9 D1 4.32 ± 0.03 b 13.64 ± 0.18 c 20.92 ± 0.10 d 50.94 ± 0.69 a
D2 4.67 ± 0.07 ab 14.35 ± 0.18 b 22.20 ± 0.32 c 50.79 ± 0.20 a
D3 4.90 ± 0.03 a 14.76 ± 0.29 ab 22.99 ± 0.32 b 50.46 ± 1.20 a
D4 5.09 ± 0.37 a 15.15 ± 0.25 a 23.64 ± 0.04 a 49.91 ± 0.18 a
Mean 4.74 14.48 22.44 50.53
Songxiangjing 1018 D1 4.79 ± 0.04 d 14.30 ± 0.12 d 22.32 ± 0.26 c 52.52 ± 0.41 a
D2 5.21 ± 0.10 c 14.80 ± 0.33 c 23.32 ± 0.30 b 52.34 ± 0.60 a
D3 5.56 ± 0.06 b 15.29 ± 0.18 b 24.26 ± 0.23 a 51.85 ± 0.51 a
D4 5.80 ± 0.13 a 15.87 ± 0.15 a 24.88 ± 0.29 a 51.29 ± 0.97 a
Mean 5.34 15.07 23.69 52.00
2024 Ningxiangjing 9 D1 4.17 ± 0.05 c 13.69 ± 0.14 c 20.85 ± 0.11 d 50.94 ± 0.25 a
D2 4.44 ± 0.06 bc 14.14 ± 0.06 b 21.86 ± 0.13 c 50.63 ± 0.23 a
D3 4.67 ± 0.06 ab 14.46 ± 0.13 a 22.62 ± 0.28 b 50.57 ± 0.47 a
D4 4.81 ± 0.24 a 14.69 ± 0.11 a 23.18 ± 0.14 a 50.45 ± 0.23 a
Mean 4.52 14.24 22.13 50.65
Songxiangjing 1018 D1 4.52 ± 0.10 d 13.90 ± 0.15 d 21.54 ± 0.16 d 52.11 ± 0.51 a
D2 4.97 ± 0.07 c 14.49 ± 0.05 c 22.65 ± 0.19 c 51.71 ± 0.16 a
D3 5.30 ± 0.09 b 15.00 ± 0.17 b 23.76 ± 0.23 b 51.39 ± 0.41 ab
D4 5.61 ± 0.11 a 15.63 ± 0.12 a 24.88 ± 0.20 a 50.72 ± 0.37 b
Mean 5.10 14.75 23.21 51.48
Year (Y) ** ** ** NS
Variety (V) ** ** ** **
Treatment (T) ** ** ** **
Y × V NS NS NS NS
Y × T NS NS NS NS
V × T NS NS NS NS
Y × V × T NS NS NS NS

Table 2. Effects of planting density on dry matter accumulation and harvest index of japonica rice.

Year Variety Treatment Dry matter (t/hm2) Harvest index
(%)
Jointing stage Heading stage Maturity stage
2023 Ningxiangjing 9 D1 4.32 ± 0.03 b 13.64 ± 0.18 c 20.92 ± 0.10 d 50.94 ± 0.69 a
D2 4.67 ± 0.07 ab 14.35 ± 0.18 b 22.20 ± 0.32 c 50.79 ± 0.20 a
D3 4.90 ± 0.03 a 14.76 ± 0.29 ab 22.99 ± 0.32 b 50.46 ± 1.20 a
D4 5.09 ± 0.37 a 15.15 ± 0.25 a 23.64 ± 0.04 a 49.91 ± 0.18 a
Mean 4.74 14.48 22.44 50.53
Songxiangjing 1018 D1 4.79 ± 0.04 d 14.30 ± 0.12 d 22.32 ± 0.26 c 52.52 ± 0.41 a
D2 5.21 ± 0.10 c 14.80 ± 0.33 c 23.32 ± 0.30 b 52.34 ± 0.60 a
D3 5.56 ± 0.06 b 15.29 ± 0.18 b 24.26 ± 0.23 a 51.85 ± 0.51 a
D4 5.80 ± 0.13 a 15.87 ± 0.15 a 24.88 ± 0.29 a 51.29 ± 0.97 a
Mean 5.34 15.07 23.69 52.00
2024 Ningxiangjing 9 D1 4.17 ± 0.05 c 13.69 ± 0.14 c 20.85 ± 0.11 d 50.94 ± 0.25 a
D2 4.44 ± 0.06 bc 14.14 ± 0.06 b 21.86 ± 0.13 c 50.63 ± 0.23 a
D3 4.67 ± 0.06 ab 14.46 ± 0.13 a 22.62 ± 0.28 b 50.57 ± 0.47 a
D4 4.81 ± 0.24 a 14.69 ± 0.11 a 23.18 ± 0.14 a 50.45 ± 0.23 a
Mean 4.52 14.24 22.13 50.65
Songxiangjing 1018 D1 4.52 ± 0.10 d 13.90 ± 0.15 d 21.54 ± 0.16 d 52.11 ± 0.51 a
D2 4.97 ± 0.07 c 14.49 ± 0.05 c 22.65 ± 0.19 c 51.71 ± 0.16 a
D3 5.30 ± 0.09 b 15.00 ± 0.17 b 23.76 ± 0.23 b 51.39 ± 0.41 ab
D4 5.61 ± 0.11 a 15.63 ± 0.12 a 24.88 ± 0.20 a 50.72 ± 0.37 b
Mean 5.10 14.75 23.21 51.48
Year (Y) ** ** ** NS
Variety (V) ** ** ** **
Treatment (T) ** ** ** **
Y × V NS NS NS NS
Y × T NS NS NS NS
V × T NS NS NS NS
Y × V × T NS NS NS NS
Table 3. Effects of planting density on leaf area index (LAI) of japonica rice.
Variety Treatment Jointing
stage
Heading stage Maturity
stage
Leaf area reduction rate per day
High-effective LAI High-effective LAI ratio (%) Effective LAI Effective LAI ratio (%) Total LAI
2023
NXJ D1 3.08 ± 0.08 b 5.65 ± 0.04 a 81.32 ± 0.36 a 6.34 ± 0.03 a 91.40 ± 0.12 a 6.94 ± 0.04 b 3.04 ± 0.05 a 0.0814 ± 0.0017 a
D2 3.18 ± 0.05 ab 5.71 ± 0.07 a 81.12 ± 0.30 a 6.42 ± 0.10 a 91.12 ± 0.48 a 7.04 ± 0.07 ab 3.12 ± 0.09 a 0.0816 ± 0.0010 a
D3 3.23 ± 0.11 ab 5.75 ± 0.17 a 80.74 ± 0.96 a 6.47 ± 0.17 a 90.89 ± 0.69 a 7.12 ± 0.13 ab 3.19 ± 0.11 a 0.0819 ± 0.0010 a
D4 3.28 ± 0.06 a 5.75 ± 0.07 a 80.02 ± 0.19 a 6.49 ± 0.11 a 90.30 ± 0.60 a 7.19 ± 0.08 a 3.23 ± 0.15 a 0.0825 ± 0.0018 a
Mean 3.20 5.72 80.80 6.43 90.93 7.07 3.14 0.0818
SXJ D1 3.13 ± 0.09 b 5.93 ± 0.07 a 83.42 ± 0.41 a 6.56 ± 0.09 a 92.24 ± 0.82 a 7.11 ± 0.05 c 3.10 ± 0.05 c 0.0802 ± 0.0008 a
D2 3.22 ± 0.04 ab 6.01 ± 0.10 a 83.24 ± 0.57 ab 6.66 ± 0.12 a 92.20 ± 0.84 a 7.22 ± 0.07 bc 3.20 ± 0.07 bc 0.0803 ± 0.0030 a
D3 3.30 ± 0.04 ab 6.05 ± 0.02 a 82.68 ± 0.43 ab 6.73 ± 0.05 a 92.03 ± 0.69 a 7.32 ± 0.03 ab 3.28 ± 0.04 ab 0.0806 ± 0.0008 a
D4 3.37 ± 0.09 a 6.07 ± 0.05 a 82.11 ± 0.50 b 6.77 ± 0.09 a 91.58 ± 0.45 a 7.39 ± 0.06 a 3.34 ± 0.03 a 0.0810 ± 0.0018 a
Mean 3.25 6.01 82.86 6.68 92.01 7.26 3.23 0.0805
2024
NXJ D1 2.97 ± 0.02 c 5.58 ± 0.04 a 80.88 ± 0.38 a 6.26 ± 0.02 b 90.83 ± 0.09 a 6.89 ± 0.02 c 2.96 ± 0.03 c 0.0820 ± 0.0010 a
D2 3.03 ± 0.02 bc 5.59 ± 0.06 a 80.44 ± 0.23 ab 6.30 ± 0.06 ab 90.58 ± 0.07 ab 6.95 ± 0.06 bc 3.01 ± 0.02 bc 0.0821 ± 0.0014 a
D3 3.08 ± 0.04 ab 5.64 ± 0.04 a 80.15 ± 0.30 b 6.35 ± 0.04 ab 90.37 ± 0.17 ab 7.03 ± 0.04 ab 3.07 ± 0.02 ab 0.0826 ± 0.0012 a
D4 3.15 ± 0.04 a 5.66 ± 0.06 a 79.88 ± 0.05 b 6.39 ± 0.07 a 90.21 ± 0.43 b 7.09 ± 0.07 a 3.10 ± 0.03 a 0.0831 ± 0.0008 a
Mean 3.06 5.62 80.34 6.33 90.50 6.99 3.03 0.0825
SXJ D1 3.06 ± 0.10 b 5.84 ± 0.06 a 83.28 ± 0.34 a 6.55 ± 0.04 b 93.41 ± 0.17 a 7.01 ± 0.04 c 2.91 ± 0.14 b 0.0805 ± 0.0033 a
D2 3.18 ± 0.09 ab 5.88 ± 0.11 a 82.41 ± 0.90 a 6.65 ± 0.11 ab 93.12 ± 1.08 a 7.14 ± 0.06 b 3.02 ± 0.08 ab 0.0807 ± 0.0006 a
D3 3.25 ± 0.10 ab 5.94 ± 0.06 a 82.06 ± 0.86 a 6.72 ± 0.08 ab 92.82 ± 0.74 a 7.24 ± 0.03 a 3.10 ± 0.05 ab 0.0813 ± 0.0013 a
D4 3.31 ± 0.08 a 6.00 ± 0.08 a 81.95 ± 0.88 a 6.78 ± 0.10 a 92.56 ± 0.91 a 7.32 ± 0.05 a 3.16 ± 0.02 a 0.0816 ± 0.0007 a
Mean 3.20 5.92 82.43 6.67 92.98 7.18 3.05 0.0810
Year (Y) ** ** NS * NS ** ** NS
Variety (V) ** ** ** ** ** ** NS *
Treatment (T) ** * ** ** NS ** ** NS
Y × V * NS NS * ** NS NS NS
Y × T NS NS NS NS NS NS NS NS
V × T NS NS NS NS NS NS NS NS
Y × V ×T NS NS NS NS NS NS NS NS

Table 3. Effects of planting density on leaf area index (LAI) of japonica rice.

Variety Treatment Jointing
stage
Heading stage Maturity
stage
Leaf area reduction rate per day
High-effective LAI High-effective LAI ratio (%) Effective LAI Effective LAI ratio (%) Total LAI
2023
NXJ D1 3.08 ± 0.08 b 5.65 ± 0.04 a 81.32 ± 0.36 a 6.34 ± 0.03 a 91.40 ± 0.12 a 6.94 ± 0.04 b 3.04 ± 0.05 a 0.0814 ± 0.0017 a
D2 3.18 ± 0.05 ab 5.71 ± 0.07 a 81.12 ± 0.30 a 6.42 ± 0.10 a 91.12 ± 0.48 a 7.04 ± 0.07 ab 3.12 ± 0.09 a 0.0816 ± 0.0010 a
D3 3.23 ± 0.11 ab 5.75 ± 0.17 a 80.74 ± 0.96 a 6.47 ± 0.17 a 90.89 ± 0.69 a 7.12 ± 0.13 ab 3.19 ± 0.11 a 0.0819 ± 0.0010 a
D4 3.28 ± 0.06 a 5.75 ± 0.07 a 80.02 ± 0.19 a 6.49 ± 0.11 a 90.30 ± 0.60 a 7.19 ± 0.08 a 3.23 ± 0.15 a 0.0825 ± 0.0018 a
Mean 3.20 5.72 80.80 6.43 90.93 7.07 3.14 0.0818
SXJ D1 3.13 ± 0.09 b 5.93 ± 0.07 a 83.42 ± 0.41 a 6.56 ± 0.09 a 92.24 ± 0.82 a 7.11 ± 0.05 c 3.10 ± 0.05 c 0.0802 ± 0.0008 a
D2 3.22 ± 0.04 ab 6.01 ± 0.10 a 83.24 ± 0.57 ab 6.66 ± 0.12 a 92.20 ± 0.84 a 7.22 ± 0.07 bc 3.20 ± 0.07 bc 0.0803 ± 0.0030 a
D3 3.30 ± 0.04 ab 6.05 ± 0.02 a 82.68 ± 0.43 ab 6.73 ± 0.05 a 92.03 ± 0.69 a 7.32 ± 0.03 ab 3.28 ± 0.04 ab 0.0806 ± 0.0008 a
D4 3.37 ± 0.09 a 6.07 ± 0.05 a 82.11 ± 0.50 b 6.77 ± 0.09 a 91.58 ± 0.45 a 7.39 ± 0.06 a 3.34 ± 0.03 a 0.0810 ± 0.0018 a
Mean 3.25 6.01 82.86 6.68 92.01 7.26 3.23 0.0805
2024
NXJ D1 2.97 ± 0.02 c 5.58 ± 0.04 a 80.88 ± 0.38 a 6.26 ± 0.02 b 90.83 ± 0.09 a 6.89 ± 0.02 c 2.96 ± 0.03 c 0.0820 ± 0.0010 a
D2 3.03 ± 0.02 bc 5.59 ± 0.06 a 80.44 ± 0.23 ab 6.30 ± 0.06 ab 90.58 ± 0.07 ab 6.95 ± 0.06 bc 3.01 ± 0.02 bc 0.0821 ± 0.0014 a
D3 3.08 ± 0.04 ab 5.64 ± 0.04 a 80.15 ± 0.30 b 6.35 ± 0.04 ab 90.37 ± 0.17 ab 7.03 ± 0.04 ab 3.07 ± 0.02 ab 0.0826 ± 0.0012 a
D4 3.15 ± 0.04 a 5.66 ± 0.06 a 79.88 ± 0.05 b 6.39 ± 0.07 a 90.21 ± 0.43 b 7.09 ± 0.07 a 3.10 ± 0.03 a 0.0831 ± 0.0008 a
Mean 3.06 5.62 80.34 6.33 90.50 6.99 3.03 0.0825
SXJ D1 3.06 ± 0.10 b 5.84 ± 0.06 a 83.28 ± 0.34 a 6.55 ± 0.04 b 93.41 ± 0.17 a 7.01 ± 0.04 c 2.91 ± 0.14 b 0.0805 ± 0.0033 a
D2 3.18 ± 0.09 ab 5.88 ± 0.11 a 82.41 ± 0.90 a 6.65 ± 0.11 ab 93.12 ± 1.08 a 7.14 ± 0.06 b 3.02 ± 0.08 ab 0.0807 ± 0.0006 a
D3 3.25 ± 0.10 ab 5.94 ± 0.06 a 82.06 ± 0.86 a 6.72 ± 0.08 ab 92.82 ± 0.74 a 7.24 ± 0.03 a 3.10 ± 0.05 ab 0.0813 ± 0.0013 a
D4 3.31 ± 0.08 a 6.00 ± 0.08 a 81.95 ± 0.88 a 6.78 ± 0.10 a 92.56 ± 0.91 a 7.32 ± 0.05 a 3.16 ± 0.02 a 0.0816 ± 0.0007 a
Mean 3.20 5.92 82.43 6.67 92.98 7.18 3.05 0.0810
Year (Y) ** ** NS * NS ** ** NS
Variety (V) ** ** ** ** ** ** NS *
Treatment (T) ** * ** ** NS ** ** NS
Y × V * NS NS * ** NS NS NS
Y × T NS NS NS NS NS NS NS NS
V × T NS NS NS NS NS NS NS NS
Y × V ×T NS NS NS NS NS NS NS NS
Table 4. Effects of planting density on plant height and top three leaves length and width of japonica rice plants.
Year Variety Treatment Plant height
(cm)
Flag leaf Second leaf from top Third leaf from top
Leaf length
(cm)
Leaf width
(cm)
Leaf length (cm) Leaf width
(cm)
Leaf length (cm) Leaf width
(cm)
2023 NXJ D1 103.00 ± 2.83 a 27.10 ± 1.00 a 1.80 ± 0.08 a 36.67 ± 2.01 a 1.77 ± 0.05 a 41.43 ± 1.98 a 1.67 ± 0.12 a
D2 102.17 ± 2.09 a 26.17 ± 0.62 a 1.77 ± 0.12 a 36.40 ± 0.91 a 1.73 ± 0.05 a 41.00 ± 1.28 a 1.63 ± 0.09 a
D3 100.83 ± 1.55 a 25.67 ± 1.39 a 1.70 ± 0.08 a 35.93 ± 1.57 a 1.67 ± 0.17 a 40.43 ± 1.14 a 1.53 ± 0.12 a
D4 97.50 ± 3.34 a 24.97 ± 1.18 a 1.63 ± 0.12 a 34.97 ± 0.52 a 1.58 ± 0.13 a 39.77 ± 0.74 a 1.43 ± 0.05 a
Mean 100.88 25.98 1.73 35.99 1.69 40.66 1.57
SXJ D1 98.00 ± 2.55 a 28.23 ± 2.05 a 1.93 ± 0.12 a 38.77 ± 0.97 a 1.90 ± 0.08 a 43.80 ± 0.43 a 1.80 ± 0.08 a
D2 97.33 ± 1.65 a 27.63 ± 2.42 a 1.90 ± 0.08 a 38.23 ± 1.32 a 1.87 ± 0.09 a 43.60 ± 1.44 a 1.77 ± 0.12 a
D3 96.17 ± 3.01 a 26.67 ± 0.74 a 1.83 ± 0.09 a 37.80 ± 1.53 a 1.83 ± 0.12 a 42.87 ± 1.93 a 1.67 ± 0.05 a
D4 94.50 ± 3.08 a 25.47 ± 2.01 a 1.77 ± 0.12 a 36.83 ± 1.07 a 1.73 ± 0.17 a 42.20 ± 1.31 a 1.60 ± 0.14 a
Mean 96.50 27.00 1.86 37.91 1.83 43.12 1.71
2024 NXJ D1 101.67 ± 2.62 a 26.60 ± 0.78 a 1.87 ± 0.21 a 36.33 ± 0.83 a 1.73 ± 0.05 a 40.10 ± 0.29 a 1.63 ± 0.05 a
D2 100.83 ± 2.25 a 26.07 ± 0.54 a 1.73 ± 0.05 a 35.73 ± 1.09 a 1.70 ± 0.08 a 39.67 ± 0.81 a 1.60 ± 0.08 a
D3 99.50 ± 1.08 a 25.30 ± 1.42 a 1.67 ± 0.12 a 34.80 ± 0.65 a 1.63 ± 0.05 a 38.43 ± 2.20 a 1.50 ± 0.08 ab
D4 97.17 ± 2.32 a 24.30 ± 1.80 a 1.60 ± 0.08 a 34.03 ± 1.22 a 1.57 ± 0.09 a 37.03 ± 1.92 a 1.37 ± 0.05 b
Mean 99.79 25.57 1.72 35.23 1.66 38.81 1.53
SXJ D1 97.67 ± 1.65 a 27.57 ± 2.11 a 1.90 ± 0.16 a 37.77 ± 2.38 a 1.87 ± 0.17 a 42.07 ± 1.20 a 1.77 ± 0.09 a
D2 96.50 ± 1.08 a 26.98 ± 1.52 a 1.87 ± 0.09 a 37.33 ± 1.77 a 1.80 ± 0.08 a 41.93 ± 1.43 a 1.70 ± 0.08 a
D3 95.00 ± 2.68 a 26.17 ± 1.54 a 1.80 ± 0.14 a 36.80 ± 1.00 a 1.73 ± 0.17 a 40.53 ± 0.62 a 1.63 ± 0.05 ab
D4 93.83 ± 3.01 a 25.13 ± 2.02 a 1.70 ± 0.08 a 36.17 ± 1.88 a 1.67 ± 0.09 a 39.87 ± 1.27 a 1.50 ± 0.08 b
Mean 95.75 26.46 1.82 37.02 1.77 41.10 1.65
Year (Y) NS NS NS NS NS ** NS
Variety (V) ** NS NS * * ** **
Treatment (T) * * ** NS NS * **
Y × V NS NS NS NS NS NS NS
Y × T NS NS NS NS NS NS NS
V × T NS NS NS NS NS NS NS
Y × V × T NS NS NS NS NS NS NS

Table 4. Effects of planting density on plant height and top three leaves length and width of japonica rice plants.

Year Variety Treatment Plant height
(cm)
Flag leaf Second leaf from top Third leaf from top
Leaf length
(cm)
Leaf width
(cm)
Leaf length (cm) Leaf width
(cm)
Leaf length (cm) Leaf width
(cm)
2023 NXJ D1 103.00 ± 2.83 a 27.10 ± 1.00 a 1.80 ± 0.08 a 36.67 ± 2.01 a 1.77 ± 0.05 a 41.43 ± 1.98 a 1.67 ± 0.12 a
D2 102.17 ± 2.09 a 26.17 ± 0.62 a 1.77 ± 0.12 a 36.40 ± 0.91 a 1.73 ± 0.05 a 41.00 ± 1.28 a 1.63 ± 0.09 a
D3 100.83 ± 1.55 a 25.67 ± 1.39 a 1.70 ± 0.08 a 35.93 ± 1.57 a 1.67 ± 0.17 a 40.43 ± 1.14 a 1.53 ± 0.12 a
D4 97.50 ± 3.34 a 24.97 ± 1.18 a 1.63 ± 0.12 a 34.97 ± 0.52 a 1.58 ± 0.13 a 39.77 ± 0.74 a 1.43 ± 0.05 a
Mean 100.88 25.98 1.73 35.99 1.69 40.66 1.57
SXJ D1 98.00 ± 2.55 a 28.23 ± 2.05 a 1.93 ± 0.12 a 38.77 ± 0.97 a 1.90 ± 0.08 a 43.80 ± 0.43 a 1.80 ± 0.08 a
D2 97.33 ± 1.65 a 27.63 ± 2.42 a 1.90 ± 0.08 a 38.23 ± 1.32 a 1.87 ± 0.09 a 43.60 ± 1.44 a 1.77 ± 0.12 a
D3 96.17 ± 3.01 a 26.67 ± 0.74 a 1.83 ± 0.09 a 37.80 ± 1.53 a 1.83 ± 0.12 a 42.87 ± 1.93 a 1.67 ± 0.05 a
D4 94.50 ± 3.08 a 25.47 ± 2.01 a 1.77 ± 0.12 a 36.83 ± 1.07 a 1.73 ± 0.17 a 42.20 ± 1.31 a 1.60 ± 0.14 a
Mean 96.50 27.00 1.86 37.91 1.83 43.12 1.71
2024 NXJ D1 101.67 ± 2.62 a 26.60 ± 0.78 a 1.87 ± 0.21 a 36.33 ± 0.83 a 1.73 ± 0.05 a 40.10 ± 0.29 a 1.63 ± 0.05 a
D2 100.83 ± 2.25 a 26.07 ± 0.54 a 1.73 ± 0.05 a 35.73 ± 1.09 a 1.70 ± 0.08 a 39.67 ± 0.81 a 1.60 ± 0.08 a
D3 99.50 ± 1.08 a 25.30 ± 1.42 a 1.67 ± 0.12 a 34.80 ± 0.65 a 1.63 ± 0.05 a 38.43 ± 2.20 a 1.50 ± 0.08 ab
D4 97.17 ± 2.32 a 24.30 ± 1.80 a 1.60 ± 0.08 a 34.03 ± 1.22 a 1.57 ± 0.09 a 37.03 ± 1.92 a 1.37 ± 0.05 b
Mean 99.79 25.57 1.72 35.23 1.66 38.81 1.53
SXJ D1 97.67 ± 1.65 a 27.57 ± 2.11 a 1.90 ± 0.16 a 37.77 ± 2.38 a 1.87 ± 0.17 a 42.07 ± 1.20 a 1.77 ± 0.09 a
D2 96.50 ± 1.08 a 26.98 ± 1.52 a 1.87 ± 0.09 a 37.33 ± 1.77 a 1.80 ± 0.08 a 41.93 ± 1.43 a 1.70 ± 0.08 a
D3 95.00 ± 2.68 a 26.17 ± 1.54 a 1.80 ± 0.14 a 36.80 ± 1.00 a 1.73 ± 0.17 a 40.53 ± 0.62 a 1.63 ± 0.05 ab
D4 93.83 ± 3.01 a 25.13 ± 2.02 a 1.70 ± 0.08 a 36.17 ± 1.88 a 1.67 ± 0.09 a 39.87 ± 1.27 a 1.50 ± 0.08 b
Mean 95.75 26.46 1.82 37.02 1.77 41.10 1.65
Year (Y) NS NS NS NS NS ** NS
Variety (V) ** NS NS * * ** **
Treatment (T) * * ** NS NS * **
Y × V NS NS NS NS NS NS NS
Y × T NS NS NS NS NS NS NS
V × T NS NS NS NS NS NS NS
Y × V × T NS NS NS NS NS NS NS
Table 5. Effects of planting density on panicle pattern of japonica rice.
Variety Treat-ment Panicle length (cm) No. of spikelets per panicle No. of filled spikelets per panicle Seed-setting rate (%) Grain density (Grain/cm) Single panicle weight (g) Axis
No. of spikelets No. of filled spikelets Seed-setting rate (%)
2023
NXJ D1 17.58 ± 0.11 a 182.86 ± 0.92 a 155.19 ± 0.73 a 84.87 ± 0.73 a 10.40 ± 0.10 a 3.80 ± 0.02 a 6.64 ± 0.08 a 6.13 ± 0.09 a 92.39 ± 2.30 a
D2 17.54 ± 0.11 a 182.08 ± 0.60 a 154.21 ± 1.34 a 84.70 ± 1.01 a 10.38 ± 0.09 a 3.77 ± 0.01 ab 6.61 ± 0.13 a 6.07 ± 0.05 ab 91.91 ± 1.60 a
D3 17.46 ± 0.09 a 180.62 ± 0.93 ab 151.92 ± 0.84 b 84.11 ± 0.09 a 10.35 ± 0.07 a 3.70 ± 0.07 bc 6.58 ± 0.07 a 6.02 ± 0.11 ab 91.44 ± 0.69 a
D4 17.34 ± 0.08 a 178.44 ± 1.60 b 149.55 ± 0.50 c 83.82 ± 0.93 a 10.29 ± 0.09 a 3.61 ± 0.01 c 6.49 ± 0.11 a 5.86 ± 0.12 b 90.21 ± 1.93 a
Mean 17.48 181.00 152.72 84.37 10.36 3.72 6.58 6.02 91.49
SXJ D1 16.46 ± 0.09 a 171.05 ± 0.57 a 151.19 ± 0.91 a 88.39 ± 0.52 a 10.39 ± 0.07 a 4.22 ± 0.02 a 6.20 ± 0.10 a 5.84 ± 0.10 a 94.20 ± 0.96 a
D2 16.44 ± 0.06 a 168.94 ± 0.88 a 148.89 ± 0.86 a 88.13 ± 0.24 a 10.28 ± 0.06 a 4.15 ± 0.01 b 6.14 ± 0.10 a 5.74 ± 0.09 ab 93.49 ± 1.34 a
D3 16.39 ± 0.07 a 165.62 ± 1.06 b 145.19 ± 1.31 b 87.67 ± 0.28 a 10.11 ± 0.04 b 4.03 ± 0.06 c 6.06 ± 0.08 a 5.63 ± 0.11 ab 92.85 ± 0.49 a
D4 16.35 ± 0.10 a 162.86 ± 1.32 c 142.27 ± 1.47 c 87.36 ± 0.58 a 9.96 ± 0.05 c 3.93 ± 0.01 d 5.98 ± 0.10 a 5.54 ± 0.05 b 92.65 ± 0.78 a
Mean 16.41 167.12 146.89 87.89 10.18 4.08 6.10 5.69 93.30
2024
NXJ D1 17.40 ± 0.18 a 175.63 ± 0.30 a 148.82 ± 0.53 a 84.74 ± 0.35 a 10.10 ± 0.10 a 3.62 ± 0.01 a 6.34 ± 0.11 a 5.80 ± 0.07 a 91.49 ± 0.67 a
D2 17.22 ± 0.14 ab 173.26 ± 0.31 b 146.59 ± 0.12 b 84.61 ± 0.20 a 10.06 ± 0.09 a 3.55 ± 0.01 b 6.30 ± 0.02 a 5.75 ± 0.05 ab 91.22 ± 0.62 a
D3 17.07 ± 0.05 bc 171.04 ± 0.09 c 144.07 ± 0.37 c 84.23 ± 0.20 a 10.02 ± 0.03 a 3.49 ± 0.01 c 6.28 ± 0.02 a 5.70 ± 0.02 ab 90.66 ± 0.17 a
D4 16.85 ± 0.08 c 167.55 ± 1.75 d 140.16 ± 0.53 d 83.67 ± 1.19 a 9.94 ± 0.11 a 3.39 ± 0.02 d 6.25 ± 0.03 a 5.66 ± 0.04 b 90.57 ± 0.90 a
Mean 17.14 171.87 144.91 84.31 10.03 3.51 6.29 5.73 90.98
SXJ D1 16.31 ± 0.02 a 165.88 ± 0.66 a 145.34 ± 0.17 a 87.62 ± 0.30 a 10.17 ± 0.05 a 4.03 ± 0.01 a 5.95 ± 0.04 a 5.49 ± 0.05 a 92.16 ± 0.53 a
D2 16.25 ± 0.03 a 163.06 ± 0.59 b 142.59 ± 0.43 b 87.44 ± 0.20 a 10.04 ± 0.05 b 3.95 ± 0.01 b 5.80 ± 0.11 ab 5.34 ± 0.03 b 92.04 ± 1.48 a
D3 16.17 ± 0.05 b 160.47 ± 0.34 c 139.89 ± 0.29 c 87.17 ± 0.17 a 9.92 ± 0.03 c 3.87 ± 0.01 c 5.70 ± 0.05 bc 5.20 ± 0.06 c 91.30 ± 1.60 a
D4 16.09 ± 0.01 c 158.45 ± 0.17 d 137.10 ± 0.28 d 86.53 ± 0.17 b 9.85 ± 0.01 c 3.78 ± 0.01 d 5.56 ± 0.09 c 5.03 ± 0.07 d 90.47 ± 0.42 a
Mean 16.21 161.97 141.23 87.19 9.99 3.91 5.75 5.26 91.49
Year (Y) ** ** ** NS ** ** ** ** NS
Variety (V) ** ** ** ** * ** ** ** NS
Treatment (T) ** ** ** ** ** ** ** ** NS
Y × V NS ** NS NS NS NS * NS NS
Y × T NS NS NS NS NS NS NS NS NS
V × T NS NS * NS ** NS ** * NS
Y × V × T NS NS * NS NS NS NS NS NS

Table 5. Effects of planting density on panicle pattern of japonica rice.

Variety Treat-ment Panicle length (cm) No. of spikelets per panicle No. of filled spikelets per panicle Seed-setting rate (%) Grain density (Grain/cm) Single panicle weight (g) Axis
No. of spikelets No. of filled spikelets Seed-setting rate (%)
2023
NXJ D1 17.58 ± 0.11 a 182.86 ± 0.92 a 155.19 ± 0.73 a 84.87 ± 0.73 a 10.40 ± 0.10 a 3.80 ± 0.02 a 6.64 ± 0.08 a 6.13 ± 0.09 a 92.39 ± 2.30 a
D2 17.54 ± 0.11 a 182.08 ± 0.60 a 154.21 ± 1.34 a 84.70 ± 1.01 a 10.38 ± 0.09 a 3.77 ± 0.01 ab 6.61 ± 0.13 a 6.07 ± 0.05 ab 91.91 ± 1.60 a
D3 17.46 ± 0.09 a 180.62 ± 0.93 ab 151.92 ± 0.84 b 84.11 ± 0.09 a 10.35 ± 0.07 a 3.70 ± 0.07 bc 6.58 ± 0.07 a 6.02 ± 0.11 ab 91.44 ± 0.69 a
D4 17.34 ± 0.08 a 178.44 ± 1.60 b 149.55 ± 0.50 c 83.82 ± 0.93 a 10.29 ± 0.09 a 3.61 ± 0.01 c 6.49 ± 0.11 a 5.86 ± 0.12 b 90.21 ± 1.93 a
Mean 17.48 181.00 152.72 84.37 10.36 3.72 6.58 6.02 91.49
SXJ D1 16.46 ± 0.09 a 171.05 ± 0.57 a 151.19 ± 0.91 a 88.39 ± 0.52 a 10.39 ± 0.07 a 4.22 ± 0.02 a 6.20 ± 0.10 a 5.84 ± 0.10 a 94.20 ± 0.96 a
D2 16.44 ± 0.06 a 168.94 ± 0.88 a 148.89 ± 0.86 a 88.13 ± 0.24 a 10.28 ± 0.06 a 4.15 ± 0.01 b 6.14 ± 0.10 a 5.74 ± 0.09 ab 93.49 ± 1.34 a
D3 16.39 ± 0.07 a 165.62 ± 1.06 b 145.19 ± 1.31 b 87.67 ± 0.28 a 10.11 ± 0.04 b 4.03 ± 0.06 c 6.06 ± 0.08 a 5.63 ± 0.11 ab 92.85 ± 0.49 a
D4 16.35 ± 0.10 a 162.86 ± 1.32 c 142.27 ± 1.47 c 87.36 ± 0.58 a 9.96 ± 0.05 c 3.93 ± 0.01 d 5.98 ± 0.10 a 5.54 ± 0.05 b 92.65 ± 0.78 a
Mean 16.41 167.12 146.89 87.89 10.18 4.08 6.10 5.69 93.30
2024
NXJ D1 17.40 ± 0.18 a 175.63 ± 0.30 a 148.82 ± 0.53 a 84.74 ± 0.35 a 10.10 ± 0.10 a 3.62 ± 0.01 a 6.34 ± 0.11 a 5.80 ± 0.07 a 91.49 ± 0.67 a
D2 17.22 ± 0.14 ab 173.26 ± 0.31 b 146.59 ± 0.12 b 84.61 ± 0.20 a 10.06 ± 0.09 a 3.55 ± 0.01 b 6.30 ± 0.02 a 5.75 ± 0.05 ab 91.22 ± 0.62 a
D3 17.07 ± 0.05 bc 171.04 ± 0.09 c 144.07 ± 0.37 c 84.23 ± 0.20 a 10.02 ± 0.03 a 3.49 ± 0.01 c 6.28 ± 0.02 a 5.70 ± 0.02 ab 90.66 ± 0.17 a
D4 16.85 ± 0.08 c 167.55 ± 1.75 d 140.16 ± 0.53 d 83.67 ± 1.19 a 9.94 ± 0.11 a 3.39 ± 0.02 d 6.25 ± 0.03 a 5.66 ± 0.04 b 90.57 ± 0.90 a
Mean 17.14 171.87 144.91 84.31 10.03 3.51 6.29 5.73 90.98
SXJ D1 16.31 ± 0.02 a 165.88 ± 0.66 a 145.34 ± 0.17 a 87.62 ± 0.30 a 10.17 ± 0.05 a 4.03 ± 0.01 a 5.95 ± 0.04 a 5.49 ± 0.05 a 92.16 ± 0.53 a
D2 16.25 ± 0.03 a 163.06 ± 0.59 b 142.59 ± 0.43 b 87.44 ± 0.20 a 10.04 ± 0.05 b 3.95 ± 0.01 b 5.80 ± 0.11 ab 5.34 ± 0.03 b 92.04 ± 1.48 a
D3 16.17 ± 0.05 b 160.47 ± 0.34 c 139.89 ± 0.29 c 87.17 ± 0.17 a 9.92 ± 0.03 c 3.87 ± 0.01 c 5.70 ± 0.05 bc 5.20 ± 0.06 c 91.30 ± 1.60 a
D4 16.09 ± 0.01 c 158.45 ± 0.17 d 137.10 ± 0.28 d 86.53 ± 0.17 b 9.85 ± 0.01 c 3.78 ± 0.01 d 5.56 ± 0.09 c 5.03 ± 0.07 d 90.47 ± 0.42 a
Mean 16.21 161.97 141.23 87.19 9.99 3.91 5.75 5.26 91.49
Year (Y) ** ** ** NS ** ** ** ** NS
Variety (V) ** ** ** ** * ** ** ** NS
Treatment (T) ** ** ** ** ** ** ** ** NS
Y × V NS ** NS NS NS NS * NS NS
Y × T NS NS NS NS NS NS NS NS NS
V × T NS NS * NS ** NS ** * NS
Y × V × T NS NS * NS NS NS NS NS NS
Fig. 1. Effects of planting density on activities of photosynthesis-related, antioxidant, and nitrogen metabolism-related enzymes in japonica rice. D1, Planting density of 16 cm × 30 cm; D2, Planting density of 14 cm × 30 cm; D3, Planting density of 12 cm × 30 cm; D4, Planting density of 10 cm × 30 cm; TPS, Tillering peak stage; JS, Jointing stage; BS, Booting stage; HS, Heading stage; FHS, Full heading stage; MFS, Mid-filling stage; RuBisCO, Ribulose-1,5-bisphosphate carboxylase; ATPase, Adenosine triphosphatase; POD, Peroxidase; SOD, Superoxide dismutase; NR, Nitrate reductase; GS, Glutamine synthetase. Data are mean ± SE (n  =  3). Different lowercase letters above bars indicate significant differences at P ≤ 0.05.

Fig. 1. Effects of planting density on activities of photosynthesis-related, antioxidant, and nitrogen metabolism-related enzymes in japonica rice. D1, Planting density of 16 cm × 30 cm; D2, Planting density of 14 cm × 30 cm; D3, Planting density of 12 cm × 30 cm; D4, Planting density of 10 cm × 30 cm; TPS, Tillering peak stage; JS, Jointing stage; BS, Booting stage; HS, Heading stage; FHS, Full heading stage; MFS, Mid-filling stage; RuBisCO, Ribulose-1,5-bisphosphate carboxylase; ATPase, Adenosine triphosphatase; POD, Peroxidase; SOD, Superoxide dismutase; NR, Nitrate reductase; GS, Glutamine synthetase. Data are mean ± SE (n  =  3). Different lowercase letters above bars indicate significant differences at P ≤ 0.05.

Fig. 2. Correlation of rice population characteristics, photosynthetic efficiency, and yield. Y, Yield; SPW, Single panicle weight; LAI, Total leaf area index at the heading stage; PH, Plant height; BIO, Dry matter accumulation; TRA, Plant translocation amount; PN, Net photosynthetic rate; SOD, Superoxide dismutase activity; POD, Peroxidase activity; RuBisCO, Ribulose-1,5-bisphosphate carboxylase activity; ATPase, Adenosine triphosphatase activity; NR, Nitrate reductase activity; GS, Glutamine synthetase activity. Data were from all replicates from two years (n = 48). *, P < 0.05.

Fig. 2. Correlation of rice population characteristics, photosynthetic efficiency, and yield. Y, Yield; SPW, Single panicle weight; LAI, Total leaf area index at the heading stage; PH, Plant height; BIO, Dry matter accumulation; TRA, Plant translocation amount; PN, Net photosynthetic rate; SOD, Superoxide dismutase activity; POD, Peroxidase activity; RuBisCO, Ribulose-1,5-bisphosphate carboxylase activity; ATPase, Adenosine triphosphatase activity; NR, Nitrate reductase activity; GS, Glutamine synthetase activity. Data were from all replicates from two years (n = 48). *, P < 0.05.

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