This study was supported by the Zhejiang Provincial Natural Science Outstanding Youth Fund Continuation Project, China (Grant No. LRG25C130002), the Innovation Program of the Chinese Academy of Agricultural Sciences (Grant No. CAAS-CSCB-202402), the Zhejiang Provincial Natural Science Foundation, China (Grant No. LD24C130001), the Biological Breeding-National Science and Technology Major Projects of China (2023ZD04066), the Central Public-interest Scientific Institution Basal Research Fund, China (Grant No. Y2025YC96), and the Agricultural Science and Technology Innovation Program, China (Grant No. CAAS-ASTIP-2021-CNRRI). We thank the Public Laboratory of China National Rice Research Institute for their technical support in GFP fluorescence observation with a laser scanning confocal microscope.