This study was supported by the National Natural
Science Foundation of China (Grant No. 42020104004), the National Key Research
and Development Program of China (Grant No. 2019YFC1803705), the Field Frontier
Program of the Institute of Soil Science, China (Grant No. ISSASIP2215), the
Agricultural Science and Technology Independent Innovation Fund Project of
Jiangsu Province, China (Grant No. CX(21)2034), the Key Research and Development
Project of Jiangsu Province, China (Grant No. BE2021717), and the Youth
Innovation Promotion Association of Chinese Academy of Sciences (Grant No.
2015250).